(A) It was $TH$ Morgan who clearly proved and defined linkage on the basis of breeding experiments in fruit flies ($Drosophila$ $melanogaster$).In $19

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(A) It was $TH$ Morgan who clearly proved and defined linkage on the basis of breeding experiments in fruit flies ($Drosophila$ $melanogaster$).In $1911$,Morgan and Castle proposed the 'chromosomal' theory of linkage,which established that genes are located on chromosomes and that linkage is a physical association of genes on the same chromosome.